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Glutathione (GSH) limits browning because it reacts with the orthoquinones produced by the enzymatic oxidation of orthodiphenols to form GRP (Grape Reaction product

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Abbreviations MSM: methyl sulphonyl methane Vit C: vitamin C GPx: glutathione peroxidise GST: glutathione transferase GR: glutathione reductase HO: heme oxygenase ROS: reactive oxygen species RNS: reactive nitrogen species NO: nitric oxide CO: carbon monoxide GSH: glutathione